Gross and net primary productivity

Energy Flow, Ecosystems and the Environment · Productivity and ecosystems · note 1 of 6

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Gross and net primary productivitySpec 5.9

In short

Gross primary productivity (GPP) is the rate at which plants fix chemical energy by photosynthesis. Net primary productivity (NPP) is what remains after the plants’ own respiration (R) is subtracted: NPP = GPP − R. NPP is the energy available to primary consumers and decomposers, and is stored as new biomass.

Gross primary productivity (GPP)
The total chemical energy fixed by photosynthesis in a given area in a given time (for example kJ m⁻² year⁻¹).
Net primary productivity (NPP)
The chemical energy remaining in plant biomass after the energy lost in respiration has been taken away. It is the energy available to the next trophic level.
NPP = GPP − R (R = energy lost in plant respiration)

Plants use some of the sugars they make in respiration, to release energy for growth, active transport and other processes. This energy is eventually lost as heat. Only the remainder is stored in new plant material (growth), which is the biomass that can be eaten by herbivores or decomposed. Productivity is measured as a rate, for example kJ m⁻² year⁻¹ for energy or g m⁻² year⁻¹ for dry mass of biomass.

Energy budget of a producer: light energy, with some not absorbed, gives gross primary production; respiration loses energy as heat and the rest is net primary production, stored in biomass that passes to primary consumers and decomposers; NPP = GPP minus R. (opens full size in a new tab)
From sunlight to biomass: NPP = GPP − R.

Calculating NPP

A field of grass fixes 12 500 kJ m⁻² year⁻¹ of energy by photosynthesis. The grass loses 4 100 kJ m⁻² year⁻¹ in respiration. Calculate the net primary productivity and express it as a percentage of GPP.

  1. NPP = GPP − R
  2. NPP = 12 500 − 4 100 = 8 400 kJ m⁻² year⁻¹
  3. Percentage = 8 400 ÷ 12 500 × 100

Answer: NPP = 8 400 kJ m⁻² year⁻¹ (67.2 % of GPP)

Maths skill:

You may be given any two of GPP, R and NPP and asked for the third. Rearrange the equation first and give the unit (kJ m⁻² year⁻¹ or g m⁻² year⁻¹).

Common mistake:

Do not say that GPP is the energy absorbed from the Sun. Most light is not absorbed, and only a small percentage of the light energy that reaches the leaf is fixed.

Written and checked against the Edexcel International A Level Biology (YBI11) specification · Updated October 2026

Frequently asked questions

What is the difference between GPP and NPP?

Gross primary productivity is the total chemical energy fixed by plants in photosynthesis. Net primary productivity is the energy left after the plants’ own respiration is subtracted (NPP = GPP − R). NPP is the energy stored as new biomass and available to the next trophic level.

Why is energy transfer between trophic levels inefficient?

Only a small fraction passes on because energy is lost in respiration as heat, in egested faeces, in excretory products such as urea and in parts that are not eaten, such as bone. These losses leave less energy available at each successive trophic level.

What is the difference between a habitat and a niche?

A habitat is the place where an organism lives. A niche is the role of the species in that habitat, including what it feeds on, when it is active, the conditions it tolerates and how it interacts with other organisms. Species in the same habitat have different niches.

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